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Multiple Choice: Chapter 8 Carriage of oxygen Question 8.1. Haemoglobin a is composed of four identical subunits b has iron at the centre of haem groups which must be in the ferrous form to bind O2 c binding of oxygen is reduced in the presence of high Pco2 because CO2 and O2 compete for the same binding sites on haemoglobin d variants (such as fetal haemoglobin) often differ from normal adult haemoglobin in the precise structure of the haem group.